Professor of Biology
Founder and CEO of The Tropical Rat Lab
Curwood Professor of Biology, 1999

J. McNaughton Rosebush Distinguished Professor, 2005

Outstanding Teacher, 2006

(920) 424-3068; adler@uwosh.edu

Academic Training:

B.S. in Biology, Washington and Lee University, 1979

Ph.D. in Biology, Boston University, 1986

Positions Held:

• Postdoctoral Research Associate, Boston University, April 1986 to April 1989.
• Lecturer, Boston University, Metropolitan College, July 1986 to August 1987.
• Postdoctoral Research Fellow in Population Sciences, July 1987 to July 1990, and Postdoctoral Research
• Associate in Population Sciences, July 1990 to July 1992 Harvard University, School of Public Health, Department of Population Sciences.
• Postdoctoral Assistant Specialist, University of California of California – Davis, Department of Zoology, January to March 1988.
• Visiting Scientist and Associate Professor, Tunghai University (Taichung, Taiwan), Department of Biology, January to June 1990.
• Postdoctoral Fellow, Smithsonian Tropical Research Institute (Panama), October 1990 to December 1993.
• Visiting Scientist, International Center for Medical Training and Research (Cali, Colombia), October to November 1993.
• Assistant Professor of Biology, University of Wisconsin – Oshkosh, Department of Biology and Microbiology, January 1994 to June 1999.
• Associate Professor of Biology, University of Wisconsin – Oshkosh, Department of Biology and Microbiology, July 1999 to June 2004.
• Professor of Biology, University of Wisconsin – Oshkosh, Department of Biology and Microbiology, July 2004 to present.

Research Interests:

Tropical forest ecology, seed predation and dispersal, ecology of tropical mammals, ecology of infectious agents.

Recent Publications:

• ADLER, G. H. 2000. Tropical tree diversity, forest structure and the demography of a frugivorous rodent. Journal of Zoology, 250: 57-74.
• Lambert, T. D., and G. H. ADLER. 2000. Microhabitat use by a tropical forest rodent, Proechimys semispinosus, in central Panama. Journal of Mammalogy, 81: 700-706.
• Mangan, S. A., and G. H. ADLER. 2000. Consumption of arbuscular mycorrhizal fungi by terrestrial and arboreal small mammals in a Panamanian cloud forest. Journal of Mammalogy, 81: 563-570.
• ADLER, G. H., and K. Kielpinski. 2000. Reproductive phenology of a tropical canopy tree, Spondias mombin. Biotropica, 32:686-692.
• ADLER, G. H. 2000. Rainforest ecosystems, animal diversity. Encyclopedia of biodiversity, 5: 1-11. Academic Press, New York.
• ADLER, G. H., M. T. Becerra, F. Prado, and B. L. Travi. 2000. Ecology of spiny rats, Proechimys canicollis, in northern Colombia. Mammalia, 64: 145-153.
• Boyett, W. D., M. E. Endries, and G. H. ADLER. 2000. Colonization and extinction dynamics of opossums on small islands in Panama. Canadian Journal of Zoology, 78: 1972-1979.
• ADLER, G. H., N. I. Suntsova, V. V. Suntsov, and S. A. Mangan. 2001. Fleas (Siphonaptera) collected from small mammals in southern Viet Nam in 1997-1998. Journal of Medical Entomology, 38: 210-213.
• Terborgh, J., L. Lopez, P. Nuñez V., M. Rao, G. Shahabuddin, G. Orihuela, M.
• Riveros, R. Ascanio, G. H. ADLER, T. D. Lambert, and L. Balbas. 2001. Ecological meltdown in predator-free forest fragments. Science, 294: 1923-1926.
• ADLER, G. H. 2001. La regulación de las poblaciones de mamíferos. Pages 329-344
• in M. R. Guariguata and G. H. Kattan (editors). Ecologia y conservación de bosques neotropicales. Editorial Libro Universitario Regional, San Jose, Costa Rica.
• Mangan, S. A., and G. H. ADLER. 2002. Seasonal dispersal of arbuscular mycorrhizal fungi by spiny rats in a Neotropical forest. Oecologia,131: 587-597.
• Travi, B. L., G. H. ADLER, M. Lozano, H. Cadena, and J. Montoya-Lerma. 2002. Impact of habitat degradation on Phlebotominae (Diptera: Psychodidae) of tropical dry forests in northern Colombia, an endemic area of visceral and cutaneous leishmaniasis. Journal of Medical Entomology, 39: 451-456.
• Travi, B. L., C. Ferro, G. H. ADLER, and J. Montoya-Lerma. 2002. Canine visceral leishmaniasis: dog infectivity to sand flies from non-endemic areas. Research in Veterinary Science, 72: 83-86.
• Travi, B. L., L. T. Arteaga, A. P. Leon, and G. H. ADLER. 2002. Susceptibility of spiny rats (Proechimys semispinosus) to Leishmania (Viannia) panamensis and Leishmania (Leishmania) chagasi. Memorias Instituto Oswaldo Cruz, 97: 887-892.
• McKee, R. C., and G. H. ADLER. 2002. Tail autotomy in the Central American spiny rat, Proechimys semispinosus. Studies on Neotropical Fauna and Environment, 37: 181-185.
• Lambert, T. D., G. H. ADLER, M. Riveros C., L. Lopez, R. Ascanio, and J. Terborgh. 2002. Rodents on tropical land-bridge islands. Journal of Zoology, 260: 179-187.
• ADLER, G. H., S. L. Davis, and A. Carvajal. 2003. Bots (Diptera: Oestridae) infesting a Neotropical forest rodent, Proechimys semispinosus (Rodentia: Echimyidae), in Panama. Journal of Parasitology, 89: 693-697.
• Suntsova, N. I., V. V. Suntsov, J. C. Beaucournu, G. H. ADLER, L. T. V. Huong, and G. V. Kuznetsov. 2003. Notes on biogeography of fleas (Insecta, Siphonaptera) of Vietnam. Pp. 65-83 in Ecological and epizootological aspects of plague in Vietnam. L. P. Korzum and V. V. Suntsov (editors). GEOS, Moscow, Russia.
• Suntsov, V. V., T. V. H. Ly, and G. H. ADLER. 2003. Distribution of rodents along a gradient of disturbance on the Tay Nguyen Plateau of southern Viet Nam. Mammalia, 67: 379-383.
• ADLER, G. H., M. T. Becerra, and B. L. Travi. 2003. Feeding success of Lutzomyia evansi (Diptera: Psychodidae) on small mammal hosts in an endemic focus of Leishmania chagasi in northern Colombia. Biomedica, 23: 396-400.
• Hoey, K. A., R. R. Wise, and G. H. ADLER. 2004. Ultrastructure of echimyid and murid rodent spines. Journal of Zoology, 263: 307-315.
• Mangan, S. A., A.-H. Eom, G. H. ADLER, J. B. Yavitt, and E. A. Herre. 2004. Diversity of arbuscular mycorrhizal fungi across a fragmented forest in Panama: insular spore communities differ from mainland communities. Oecologia, 141: 687-700.
• Endries, M. E., and G. H. ADLER. 2004. Spacing patterns of a tropical forest rodent, the spiny rat (Proechimys semispinosus), in Panama. Journal of Zoology, 265: 147-155.
• Carvajal, A., and G. H. ADLER. 2005. Biogeography of mammals on tropical Pacific islands. Journal of Biogeography, 32: 1561-1569.
• ADLER, G. H., A. Carvajal, S. W. Brewer, and S. L. Davis. 2006. First record of Didelphis albiventris (Didelphimorphia: Didelphidae) from Paracou, French Guiana. Mammalia, 2006: 319-320.
• ADLER, G. H., and T. D. Lambert. 2008. Spatial and temporal variation in fruiting phenology of two species of palms in isolated stands. Plant Species Biology, 23: 9-17.
• Oaks, J. R., J. M. Daul, and G. H. ADLER. 2008. Life span of a tropical forest rodent, Proechimys semispinosus. Journal of Mammalogy, 89: 904-908.
• ADLER, G. H. 2008. Resource limitation of insular animals: causes and consequences. Pages 322-333 in W. Carson and S. Schnitzer, Tropical Forest Ecology. Blackwell.
• Carvajal, A., and G. H. ADLER. 2008. Seed dispersal and predation by Proechimys semispinosus and Sciurus granatensis in gaps and understorey in central Panama. Journal of Tropical Ecology, 24: 485-492.
• Flagel, D., G. H. ADLER, and T. D. Lambert. 2009. Influence of seed height on removal rates by rodents in central Panama. Mammalia, 73: 76-77.
• Kilgore, A., T. D. Lambert, and G. H. ADLER. 2009. Lianas influence fruit and seed use by rodents in a tropical forest. Tropical Ecology, 51:265-271.
• 86. ADLER, G. H. 2010. Spacing patterns and social mating systems of echimyid rodents. Journal of Mammalogy, 92: 31-38.
• Lambert, T. D., M. K. Halsey, J. W. Dittel, S. A. Mangan, E. Delfosse, G. H. ADLER, and S. A. Schnitzer. 2010. First record of Alston’s woolly mouse opossum (Micoureus alstoni) from the canal area of central Panama. Mammalia, 75: 107-109.

Graduated Master's Students:

• Donna M. Charley. 1996. Density dependence in the tropics: a statistical analysis of Proechimys semispinosus (spiny rats) populations.
• Current position: Senior Lecturer, University of Wisconsin – Oshkosh, Department of Biology and Microbiology, Oshkosh, WI.
• Kelley A. Hoey. 1999. External spine morphology of the genus Proechimys (Rodentia: Echimyidae) as revealed by scanning electron microscopy.
• Current position: Electron microscopist, Mayo Foundation, Rochester, MN.
• Thomas D. Lambert. 1999. The effects of tropical forest fragmentation on rodent community structure.
• Ph.D., University of Toronto.
• Current position: Assistant Professor, Frostburg State University, Department of Biology, Frostburg, MD.
• Scott A. Mangan. 1999. The consumption of arbuscular mycorrhizal fungi by Neotropical rodents.
• Ph.D., Indiana University
• Current position: Postdoctoral Research Fellow, University of Wisconsin – Milwaukee, Department of Biology, Milwaukee, WI.
• Mark J. Endries. 2000. Modeling spacing patterns of a tropical forest rodent using radiotelemetry and geographic information systems (GIS).
• Current position: Geographical Information Systems Specialist, U. S. Fish and Wildlife Service, Asheville Ecological Services Field Office, Asheville, NC.
• Shannon L. Davis. 2001. Environmental variation and the demography of a tropical forest rodent.
• Current positions: Ph.D. candidate, University of Wisconsin – Milwaukee, Department of Biology, Milwaukee, WI, and Lecturer, Department of Biology and Department of Environmental Studies, University of Wisconsin –Oshkosh, Oshkosh, WI.
• William D. Boyett. 2001. Habitat relations of rodents in the Hualapai Mountains of northwestern Arizona.
• Michael J. Lohre. 2003. Habitat associations of the whip-poor-will (Caprimulgus vociferus) in central Wisconsin: implications for conservation.
• Alejandra Carvajal. 2006. Seed dispersal and predation by Neotropical forest rodents.

Current Master's Students:

• Joshua Demski. Habitat associations of two color morphs of the red-backed salamander in northern Wisconsin.
• Jacob W. Dittel. The role of rodents in affecting forest regeneration: seed dispersal and seed predation.
• Annette Ireland. Helminth parasitism in relation to the demography of a Neotropical forest rodent.
• Julie Widholm. Thermoregulation and habitat preference in two species of tropical rodents.
• Ashley M. Winker. Parasite loads of two species of Neotropical forest rodents in relation to microhabitat structure.